; Broady, P.A. Zieger, S.; Seoane, S.; et al. Ditylum brightwellii Ditylum sol unclassified Ditylum Helicotheca Helicotheca tamesis unclassified Helicotheca Lithodesmioides Lithodesmioides polymorpha Lithodesmium Lithodesmium intricatum Lithodesmium undulatum Lithodesmium variabile unclassified Lithodesmium environmental samples uncultured Ditylum (1984). Author: Stefanidou, Natassa; Genitsaris, Savvas; Lopez-Bautista, Juan; Sommer, Ulrich; Moustaka-Gouni, Maria Source: Oecologia 2019 v.191 no.2 pp. Lithodesmiales. Free and Open Access to Biodiversity Data. New Zealand inventory of biodiversity: 3. In Puget Sound, WA, D. brightwellii is composed of two metapopulations that are defined by DNA sequence dif-ferences in the ribosomal internal transcribed spacers (ITS) [30,31]. Schizochytrium sp. ; Cassie Cooper, V.; Chang, F.H. Aracne: Roma. The phred quality scores of our sequences derived from single-cell material were similar to phred scores of our own, published sequences obtained from cultures grown in our laboratory. The simplest type of sexual reproduction between gametes that are similar in size and shape. (USA). California Academy of Sciences, On-line Version. Academic Press: San Diego, CA [etc.] Lithodesmiaceae. Le phytoplancton marin du Liban (Méditerranée orientale): biologie, biodiversité, biogéographie. Les Diatomacées (suite) (Le Genre Chaetoceros excepté). The sequential wall formation in the centric diatom,Ditylum brightwellii (West) Grunow, is described. Instead of being classified as male and female cells, organisms of different mating types (i.e., different size and shape of gamete) can fuse to form a zygote (Botany Dictionary 2002). Classification kingdom Chromista phylum ... Ditylum brightwellii Name Synonyms Triceratium brightwellii T.West, 1860 Homonyms Ditylum brightwellii (T.West) Grunow, 1885 Common names 布氏雙尾藻 in language. Silicification of the new valve starts from the central labiate process area prior to the completion of cytokinesis, and the developing valve grows in a centrifugal direction. (2012). (2011). Ditylum brightwellii Taxonomic Serial No. Mediophyceae. UniParc. (Length ~40 – ~300 µm) (Diameter ~14 – ~120 µm) Ditylum are solitary cells that are sometimes found in small chains. Hong Kong University Press, Hong Kong. [On the morphology of Ditylum brightwellii (Bacillariophyta)]. Infraspecies recognized by EOL Dynamic Hierarchy 0.9 and GBIF classification. We examined the effect of iron (Fe) and Fe-light (Fe-L) co-limitation on cellular silica (BSi), carbon (C) and nitrogen (N) in two marine diatoms, the small oceanic diatom Thalassiosira oceanica and the large coastal species Ditylum brightwellii.We showed that C and N per cell tend to decrease with increasing Fe limitation (i.e. Ditylum brightwellii. Canterford, G. S. 1980. In: Morton B, editor. 95: 210 pp. Muylaert, K.; Sabbe, K. (1999). Spectral Characterization of Eight Marine Phytoplankton Phyla and Assessing a Pigment-Based Taxonomic Discriminant Analysis for the in-Situ Classification of Phytoplankton Blooms. ... No drawings available for this family. Zieger, S.; Seoane, S.; et al. Curated hierarchies for Ditylum brightwellii var. (2012). The pervalvar axis can be 2-10 times the valve diameter, thus living cells are usually seen in girdle view. Thus far, eight different diatom viruses infecting the genera Rhizosolenia and Chaetoceros have been isolated and characterized to different extents. Phylum Ochrophyta: brown and golden-brown algae, diatoms, silicoflagellates, and kin, in: Gordon, D.P. The 21-day NOEC and LOEC values for freshwater algae ranged from 0.6 to 1.6 mg Co/L (WHO CICAD, 2006). Ditylum brightwellii belongs to order Lithodesmiales, which is a sister group of Thalassiosirales based on SSU rRNA (Medlin and Kaczmarska, 2004). Scale bars = 20 μm. Centric. Recife: SUDENE,318,1990. The sequential wall formation in the centric diatom, Ditylum brightwellii (West) Grunow, is described. ; Broady, P.A. Muller, Y. Spectral Characterization of Eight Marine Phytoplankton Phyla and Assessing a Pigment-Based Taxonomic Discriminant Analysis for the in-Situ Classification of Phytoplankton Blooms. Diatom. 60, Issue. Diatoms are one of the most significant primary producers in the ocean, and the importance of viruses as a potential source of mortality for diatoms has recently been recognized. 114-163. (2008). The objective of the present study was to assess size and morphological changes in response to temperature in the marine diatom, Ditylum brightwellii (West) Grunow, using ecological, morphological and physiological approaches. Here, we show that gene flow between distant populations of the globally distributed, bloom-forming diatom species Pseudo-nitzschia pungens … In terms of resource acquisition (nutrients, light), low availability usually favours smaller phytoplankton, which exhibit higher acquisition and growth rates at low resource availability (“affinity”). pp. ... Odontella aurita, and Ditylum brightwellii . Classification. It is a unicellular photosynthetic autotroph that has the ability to divide rapidly and contribute to spring phytoplankton blooms. Kingdoms Bacteria, Protozoa, Chromista, Plantae, Fungi. ( diatoms) Having radial symmetry, i.e., cell is shaped like a coin or a tuna can or a soup can. These web pages are currently under construction and expansion. However, the three D. brightwellii strains formed a distinct grouping based on CCM genetic structure, driven by diversification of CAs within this species ( … Two EC50 values were available for marine species with a similar range of 0.3 mg Co/L for Ditylum brightwellii and 10.2 mg Co/L for Nitzschia closterium (WHO CICAD, 2006). Referentiematrices en Ecotoopoppervlaktes Annex bij de Evaluatiemethodiek Schelde-estuarium Studie naar “Ecotoopoppervlaktes en intactness index”. Swedish Taxonomic Database. (2013). Produce resting spores. Spring phytoplankton assemblages in and around the maximum turbidity zone of the estuaries of the Elbe (Germany), the Schelde (Belgium/The Netherlands) and the Gironde (France). Ditylum brightwellii [saltwater diatom] Saved by emma klee. Seung Won Jung, Seok Jae Youn, Hyeon Ho Shin, Suk Min Yun, Jang-Seo Ki, Jin Hwan Lee, Effect of temperature on changes in size and morphology of the marine diatom, Ditylum brightwellii (West) Grunow (Bacillariophyceae), Estuarine, Coastal and Shelf Science, 10.1016/j.ecss.2013.05.007, 135, (128-136), (2013). The signature of unequalgene flow among populations suggested that D. brightwellii exhibited … D. brightwellii. -. Hällfors, G. (2004). For freshwater invertebrates, acute LC50s (24–96 h) range from 1.1 to 239 mg/L. Numerous, small chloroplasts and a long central spine. (2013). Disclaimer: ITIS taxonomy is based on the latest scientific consensus available, and is provided as a general reference source for interested parties. General. European Marine Sites, Datasets, Species and Distribution - MarBEF Integrated Data System (MarIDaS) Ditylum brightwellii var. ring populations of the common coastal diatom Ditylum brightwellii (T. West) Grunow in van Heurk, which has a wide-spread coastal and estuarine distribution. For recent examples regarding uptake at low nutrient concentrations, see Edwards et al. Diatoms possess a silicon-rich cell wall called the frustule, which is an important trait for taxonomic classification [26,27]. Family. Krayesky, D. M.; Meave, D. C.; Zamudio, E.; Norris, E.; Fredericq, S.; Tunnell, J. UniProtKB. (1988). Aug 13, 2019 - This Pin was discovered by Patricia Thomas. Environmental hazard assessments using diatoms have been well documented; however, their molecular toxicology has not been sufficiently studied. Mar 31, 2018 - This Pin was discovered by Hans de Vries. (2011) and Litchman et al. [Ruiyu] (ed.). (2002). Diatoms are sensitive indicators of water quality, and hence used for environmental hazard assessments; however, their toxicogenomic studies have been insufficiently attempted. 1, pp. Journal of the Marine Biological Association of the United Kingdom. 66(3): 531-610. (2002). Classification route species: (algal) Classification description: Very ecotoxic in the aquatic environment Classification key study: SPECIES: Ditylum brightwellii Diatom TYPE OF EXPOSURE: static DURATION: 5 d ENDPOINT: EC50 growth VALUE: T 100 ug/l REFERENCE SOURCE: Ref No: 6405. 114-163. Ditylum brightwellii: (a) single cell with outer parts of bilabiate process; (b) cell with resting spore; (c) valve view. [On the morphology of Ditylum brightwellii (Bacillariophyta)]. The silica deposition vesicle is formed by the coalescence of small vesicles. Ditylum brightwellii is very large (40 000–150 000 μm 3) and characterized by elongate cylindrical cells with a low S:V ratio (0.10–0.20 μm −1) and a moderate SI (12–23). Botanicheskii Zhurnal, 69(5): 671-674, 1 pl. Classification / Names Common names | Synonyms | CoL | ITIS | WoRMS Coscinodiscophyceae | Lithodesmidales | Lithodesmiaceae. Help. Integrated Taxonomic Information System (ITIS). Diatoms (Bacillariophyta) of the Gulf of Mexico. Approximately 200,000 diatom species are thought to exist and yet the underlying processes of speciation in diatoms are unknown. Ditylum brightwellii form siliceous ooze s. … Ditylum brightwellii (T. West) Grunow in Van Heurck is now a model organism for population genetic studies of marine phytoplankton, but observations of sexual reproduction in this species are sparse and there are controversial aspects to the identification of its gametes. -. This organism was identified by 16S rRNA sequencing as a strain with close affinities to Pseudoalteromonas peptidysin . 2010).It reproduces sexually by formation of Close Isogamy. ISBN 978-88-548-4243-4. (urn:lsid:marinespecies.org:taxname:149023), The webpage text is licensed under a Creative Commons Attribution 4.0 License, (urn:lsid:marinespecies.org:taxname:149023), Intergovernmental Oceanographic Commission (IOC) of UNESCO. Analytical flow cytometry (FCM) is well suited for the analysis of phytoplankton communities in fresh and sea waters. Phytoplankton monitoring in the Western Isles region of the Bay of Fundy during 1997-98. Helicotheca tamesis: twisted chain. (1986). Syst. Ditylum brightwellii. Species Description. A check-list of the freshwater, brackish and marine diatoms of the British Isles and adjoining coastal waters. Hartley, B., Ross, R. & Williams, D.M. Thomas, M. L. H. (1983). Attention was directed particularly at the origin of the Labiate Process Apparatus (LPA). The 5-day EC50 for cobalt based on growth of the marine diatom Ditylum brightwellii was 0.3 mg/L. Marine and coastal systems of the Quoddy Region, New Brunswick. Dyntaxa. Genus:Ditylum | Diatom. K morfologii Ditylum brightwellii (Bacillariophyta). Ditylum brightwellii Imported Taxonomic identifier i: 49249 : Taxonomic lineage i › Eukaryota › Sar › Stramenopiles › Ochrophyta › Bacillariophyta › Mediophyceae › Lithodesmiophycidae › Lithodesmiales › Lithodesmiaceae › Ditylum. 01, p. 243. (2015), and for light utilization at low intensities Edwards et al. Belongs to the actin family. Description. Twopopulations shared ITSI sequence type 1 and two shared ITSI sequence type 2,indicating differences in relatedness among populations. ; Nelson, W.A. 7. striata shrubsolei Example of cytometry scanned profiles and images : diatom species Ph.D. Thesis of Simon Bonato 2011-2014 Its morphometric traits showed the same response to light as the average community traits (Fig. Harper, M.A. Accessed at www.dyntaxa.se [15-01-2013]. Lam CWY. Identifying marine phytoplankton. Environmental Science & Technology 2018., DOI: 10.1021/acs.est.8b04528. your own Pins on Pinterest Ditylum brightwellii (T. West) Grunow in Van Heurck is now a model organism for population genetic studies of marine phytoplankton, but observations of sexual reproduction in this species are sparse and there are controversial aspects to the identification of its gametes. brightwellii has two distinct cell morphologies: prism (large-sized cell) and cylinder types (small-sized cell). Bengt Karlson (2008) Bearbetad lista över encelliga organismer i marin miljö. Starred values indicate that specific growth rates were measured at these concentrations but not elemental composi-tion. Environmental Science & Technology 2018., DOI: 10.1021/acs.est.8b04528. The simplest type of sexual reproduction between gametes that are similar in size and shape. Monitor Taskforce Publication Series, 2013-01. A taxonomic guide to some common marine phytoplankton. Atlas. Classification route species: (algal) Classification description: Very ecotoxic in the aquatic environment Classification key study: SPECIES: Ditylum brightwellii Diatom TYPE OF EXPOSURE: static DURATION: 5 d ENDPOINT: EC50 growth VALUE: T 100 ug/l REFERENCE SOURCE: Ref No: 6405. (Ed.). K morfologii Ditylum brightwellii (Bacillariophyta). Phylum Ochrophyta: brown and golden-brown algae, diatoms, silicoflagellates, and kin, in: Gordon, D.P. Mar. Ditylum brightwellii (T. West) Grunow. Biopress Ltd. Bristol. 1-195. Oxygen is a particularly important environmental factor for cell growth and DHA synthesis. Sequence archive. On the contrary, larger phytoplankton have better storage capacities, as indicated by higher ratios betw… Catalogue of Diatom Names. Faune et flore du littoral du Nord, du Pas-de-Calais et de la Belgique: inventaire. XV, 858 pp. K morfologii Ditylum brightwellii (Bacillariophyta). Abstract. Ducaju & Cie., Anvers., This service is powered by LifeWatch Belgium, http://www.sciencedirect.com/science/book/9780126930184, http://helcom.fi/Lists/Publications/BSEP95.pdf, http://www.vliz.be/imisdocs/publications/145561.pdf, http://researcharchive.calacademy.org/research/diatoms/names/index.asp, To Biodiversity Heritage Library (29 publications), To Biodiversity Heritage Library (3 publications), To GenBank (203 nucleotides; 46 proteins). Horner, R. A. Because genetic subdivision within species can reveal potential speciation mechanisms, we examined genetic differentiation and patterns of gene flow among four populations of the diatom Ditylum brightwellii. datasets have provided data to the NBN Atlas for this species.. Browse the list of datasets and find organisations you can join if you are interested in participating in a survey for species like Ditylum brightwellii (T.West) Grunow, 1885 Botanicheskii Zhurnal, 69(5): 671-674, 1 pl. Listan följer Algaebase (http://www.algaebase.org) och har bearbetats av Bengt Karlson (SMHI). Biopress Ltd. Bristol. 1-195. : 2928 (Download Help) Ditylum brightwellii TSN 2928 Taxonomy and Nomenclature Kingdom: Chromista : Taxonomic Rank: Species : Synonym(s): Common Name(s): Taxonomic Status: Current Standing: accepted … NIOZ: Yerseke. 35 pp. (2011). 461-474 ISSN: (2015, 2016) and Schwaderer et al. by Rebecca Sellsted. Genus-Ditylum. Jul 26, 2019 - This Pin was discovered by Linda M Fox. brightwellii. Updated 2011-09-19. Checklist of Baltic Sea Phytoplankton Species (including some heterotrophic protistan groups). x; UniProtKB. Electivity index (Ei) for the (plurialgal) selective feeding experiment, using a mixed assemblage of four prey (Ditylum brightwellii, Gymnodinium sp., Phaeodactylum tricornutum and Rhodomonas baltica) at three total prey concentrations (low: 56.1 ± 3.39, inter: 454.7 ± 12.99 and high: 1107.9 ± 237.08 µg C L −1). [Coastal fauna and flora of the Nord, Pas-de-Calais and Belgium: inventory]. Huntsman Marine Laboratory, St. Andrews, N.B. 68: p. Martin, J. L.; LeGresley, M. M. ; Strain, P. M. (2001). (2011). A protocol for a single-cell PCR of diatoms from fixed samples: method validation using Ditylum brightwellii (T. West) Grunow, Diatom Res., 2011, vol. Ditylum. Photosynthetic. ; Cassie Cooper, V.; Chang, F.H. Form recognized by GBIF classification. (2012). (1997). Canadian Technical Report of Fisheries and Aquatic Sciences 2349. 4: 1-85. Protein knowledgebase. Discover (and save!) The Ocean Biogeographic Information System (OBIS). Ditylum brightwellii Asterionellopsis glacialis Guinardia delicatula Rhizosolenia imbricata var. Ditylum J.W.Bailey ex L.W.Bailey, 1861. 3N–P; Table I). Order: Biddulphiales. This approach was applied to the diatom Ditylum brightwellii to assess the quality of recovered sequences. (2004). (2012). Sexual reproduction by Close Isogamy. Shape. Tomas, C.R. We’re sorry, but GBIF doesn’t work properly without JavaScript enabled. Bacillariophyceae. European Marine Sites, Datasets, Species and Distribution - MarBEF Integrated Data System (MarIDaS) 26, no. The role of geographic isolation in marine microbial speciation is hotly debated because of the high dispersal potential and large population sizes of planktonic microorganisms and the apparent lack of strong dispersal barriers in the open sea. Ditylum. your own Pins on Pinterest Microplankton de la Mer Flamande: 2. Journal of the Marine Biological Association of the United Kingdom. 66(3): 531-610. Commission Régionale de Biologie Région Nord Pas-de-Calais: France. 307 pp. Genus. Van Heurck, H. (1880). We suggest you upgrade to a modern browser. Click on the illustration to enlarge. pp. Gulf of Mexico origin, waters, and biota. 1: 155-186. Baltic Sea Environment Proceedings. No. Description: Found as single cells or short chains. (1986). Formation and Regeneration of Abnormal Cells of the Marine Diatom Ditylum Brightwellii (West) Grunow.Journal of the Marine Biological Association of the United Kingdom, Vol. Lithodesmium undulatum: (a) short chain; (b) valve with central bilabiate process and depressions across the corners. J. Asian Marine Biology 6. pp 5-18. 2017-09-14 18:27:12 Lisa Karlson - Deleted media: Ditylum brightwellii_7.jpg ; 2017-09-14 18:25:21 Lisa Karlson - Updated media metadata for Ditylum brightwellii_7.jpg ; 2017-09-14 18:24:45 Lisa Karlson - Added media: Ditylum brightwellii_7.jpg ; 2013-05-14 07:30:45 Regina Hansen - Updated media metadata for Ditylum brightwellii_6.jpg Horner, R. A. (Ed.) Fourtanier, E. & Kociolek, J. P. (compilers). Family & Domains i Sequence similarities i. Kingdom-Protista. Dzhafarova, S.K. Ditylum brightwellii: COPEPEDIA is an in-development project. Analytical flow cytometry (FCM) is well suited for the analysis of phytoplankton communities in fresh and sea waters. Ditylum brightwellii is a species of diatoms in the family Lithodesmiaceae. (Ed.) Harper, M.A. Higher Classification: > Kingdom Chromista > Subkingdom Harosa > Infrakingdom Heterokonta > Phylum Ochrophyta > Subphylum Khakista > Class Bacillariophyceae > Subclass Coscinodiscophycidae > Superorder Lithodesmiophycanae > Order Lithodesmiales > Family Lithodesmiaceae > Genus Ditylum Ditylum brightwellii (T.West) Grunow, 1885 Rank: Species However, it is not a legal authority for statutory or regulatory purposes. Discover (and save!) Centric diatom. Discover (and save!) Common West Coast species:D. brightwellii. Meunier, A. Ditylum brightwellii cells are primarily triangular, though often with rounded corners, giving the cells a cylindrical appearance (Figures 1 & 2). Neostreptotheca subindica: chain in girdle view. Accessed at www.dyntaxa.se [15-01-2013]. [On the morphology of Ditylum brightwellii (Bacillariophyta)]. (1984). D. brightwellii has two distinct cell morphologies: prism (large-sized cell) and cylinder types (small-sized cell). For freshwater invertebrates, acute LC50s (24–96 h) range from 1.1 to 239 mg/L. [ About COPEPEDIA] Order. 182 pp. (1915). Kingdoms Bacteria, Protozoa, Chromista, Plantae, Fungi. California Academy of Sciences, On-line Version. During a bacterial survey of the Huon Estuary in southern Tasmania, Australia, we isolated a yellow-pigmented Pseudoalteromonas strain (class Proteobacteria , gamma subdivision), designated strain Y, that had potent algicidal effects on harmful algal bloom species. 149023  http://creativecommons.org/licenses/by-nc/4.0/, http://creativecommons.org/licenses/by-nc-nd/4.0/, http://creativecommons.org/publicdomain/zero/1.0/, http://creativecommons.org/licenses/by-sa/4.0/, Ditylum brightwellii (T.West) Grunow, 1885. your own Pins on Pinterest In our current study we applied both spectroscopy methods to study chemical variations within individual cells of the diatom Ditylum brightwellii during the course of growth phases. ; Nelson, W.A. is a promising strain for the production of docosahexaenoic acid (DHA)-rich oil and biodiesel, and has been widely used in the food additive and bioenergy industries. A checklist of marine fauna and flora of the Bay of Fundy. Environment: milieu / climate zone / depth range / … Botanicheskii Zhurnal, 69(5): 671-674, 1 pl. Noctiluca scintillans is an athecate heterotrophic dinoflagellate that feeds by phagotrophy (Kraberg et al. Phytoplankton characteristics of Tolo Harbour. (2009). Two numerically dominant ITSI sequence types were detected and their distribution among isolates from genetically distinct populations was significantly different. Ditylum brightwelli is a species of cosmopolitan marine centric diatoms. From one view they are rectangular but … Updated 2011-09-19. Checklist of marine biota of China seas. Our website has detected that you are using an outdated insecure browser that will prevent you from using the site. Family: Lithodesmisceae. Species. Liu, J.Y. A check-list of the freshwater, brackish and marine diatoms of the British Isles and adjoining coastal waters. Canadian Special Publication of Fisheries and Aquatic Sciences. 64:1-306. Catalogue of Diatom Names. It is native to Paraná, Sergipe, Rio Grande do Sul, Bahia, Maranhao, São Paulo, Alagoas, Pernambuco, Pará, Espirito Santo, Amapá, Rio Grande do Norte, and Rio de Janeiro. Maris, T.; Beauchard, O.; Van Damme, S.; Van den Bergh, E.; Wijnhoven, S.; Meire, P. (2013). Help pages, FAQs, UniProtKB manual, documents, news archive and Biocuration projects. A taxonomic guide to some common marine phytoplankton. In our current study we applied both spectroscopy methods to study chemical variations within individual cells of the diatom Ditylum brightwellii during the course of growth phases. 43-49. doi 10.1080/0269249X.2011.573703 CrossRef Google Scholar Lakkis, S. (2011). Global Biodiversity Information Facility. Hartley, B., Ross, R. & Williams, D.M. Ditylum brightwellii (T.West) Grunow, 1885. Ditylum brightwellii Imported Taxonomic identifier i: 49249 : Taxonomic lineage i › Eukaryota › Sar › Stramenopiles › Ochrophyta › Bacillariophyta › Mediophyceae › Lithodesmiophycidae › Lithodesmiales › Lithodesmiaceae › Ditylum. There are no common names associated with this taxon. In the coastal waters of South Korea, the prism … The silica deposition vesicle is formed by the coalescence of small vesicles. Belongs to the actin family. Hayez, imprimeur de l'Académie royale de Belgique: Bruxelles. 293 pp. Dyntaxa. Dzhafarova, S.K. brightwellii. Cells of the centric diatomDitylum brightwellii were filmed undergoing cell division and valve secretion, and were fixed for transmission electron microscopy. Linkletter, L. E. (1977). Swedish Taxonomic Database. 22: 133-149. Catálogo das diatomáceas (Bacillariophyceae) da plataforma continental de Pernambuco. Fourtanier, E. & Kociolek, J. P. (compilers). Summary. Close. China Science Press. 1267 pp. New Zealand inventory of biodiversity: 3. The objective of the present study was to assess size and morphological changes in response to temperature in the marine diatom, Ditylum brightwellii (West) Grunow, using ecological, morphological and physiological approaches.D. Family & Domains i Sequence similarities i. ISBN 0-12-693018-X. Mémoires du Musée Royal d'Histoire Naturelle de Belgique = Verhandelingen van het Koninklijk Natuurhistorisch Museum van België, VII(3). Synopsis des Diatomées de Belgique. Common Names. & Ho KC. Toxicity to algae Growth inhibition EC50 - Ditylum brightwellii - 0,01 mg/l - 5 d 12.2 Persistence and degradability No data available 12.3 Bioaccumulative potential Bioaccumulation Pimephales promelas (fa thead minnow) - 0,50 µg/l Bioconcentration factor (B CF): 5.680 12.4 Mobility in soil No data available 12.5 Results of PBT and vPvB assessment

ditylum brightwellii classification

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